In this version, Remy cannot sniff (though he can skitter and look around by pressing the up button of the d-pad), run on a ball (though he can jump on a balloon or jump from bouncy surfaces), distract enemies by throwing them explosives, and use an umbrella to glide. Enemies include many insects, amphibians, rodents, and "flyers" such as bats and crows, and also bosses like Skinner, a giant sewer turtle and giant crows, who try without delay to stop him. About to open their new restaurant Le Ratatouille, Remy's recipes are stolen and he, with the help of his colony (including one teen rat who always wants to challenge him to race with him) must find the recipes by exploring Linguini's Apartment, the Sewers, a nearby Marketplace, the Rooftops, and (Skinner's) Kitchen.
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The PSP version's plot is after the events that occurred before. Remy has limited lives, can glide and collect things to help complete the level and passwords when completing levels. In this version, it only uses 2D graphics (side view and top view). Now it is up to Remy, Linguini, and the chef Colette to cook for many people, even the critic Ego. However, they all leave after finding out about Remy. After that, the food critic Anton Ego, also known as the "Grim Eater," has arrived at Gusteau's for a review one that will be very important to the cooks. Later, Remy helps his colony steal prized foods at the market. Skinner catches Remy, and another chase is on. The next day Remy helps Linguini cook the food for the customers while he helps his colony, which he reunites with, in the meantime. After that Remy joins Linguini and helps him with what he is forced to do for Skinner, the chef. Remy drops in and fixes the soup, but Linguini witnesses him, thus beginning a chase outside with Linguini on pursuit. After Remy successfully escapes the shotgun-wielding woman, he is caught in the rapids of the sewers, and he wakes up in front of Gusteau's restaurant, where a small ghost of Gusteau guides Remy to the top of the restaurant, where the rat witnesses the garbage boy, Linguini, attempting to fix the soup he accidentally spilt by scrambling in a bunch of random ingredients. After the task, the old lady living in the farm catches Remy and Emile, alerting the colony, thus forcing them to escape. On the trek he is taught the basic skills he will need to know so that he can accomplish what he faces later on. Remy goes off with his brother Emile to retrieve apple cores for his father. Similar to the plot of the movie, Remy begins in the farm setting in which the film starts as well. Several of the actors from the film voice their characters in the game.
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